Melanis (28 species) is a mostly South American genus, with 3 of the species found north of Panama. These are dark butterflies with lighter markings on or near the apices of all four wings.
M. smithiae has markings as shown, but these are variable, and for example the four red spots are sometimes absent.
The known life histories of Melanis species involve larvae developing on trees in the family Fabaceae. Adults nectar on a variety of shrubs and trees, including Orange and Inga.
Melanis smithiae has been reported from Colombia, Venezuela, Suriname, Brazil, Ecuador, Peru, Bolivia, and Paraguay.
Photo location: Novo Airão, Brazil.
American Insects site